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Wikipedia says:

"evidence of arthropod boreholes in Prototaxites has been found from the early and late Devonian, suggesting the organism survived the stress of boring for many millions of years.[30] Intriguingly, boreholes appeared in Prototaxites long before plants developed a structurally equivalent woody stem, and it is possible that the borers transferred to plants when these evolved"

and for Arthropods:

"The evolutionary ancestry of arthropods dates back to the Cambrian period."

which is about 100 million years before the Devonian

also:

"The oldest known arachnid is the trigonotarbid Palaeotarbus jerami, from about 420 million years ago in the Silurian period.[82][Note 3] Attercopus fimbriunguis, from 386 million years ago in the Devonian period, bears the earliest known silk-producing spigots"

so there could have been all sorts of weird and wonderful bugs burrowing into Prototaxites I guess



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